Ph.D. in Biological Sciences. The University of Texas, Austin, Texas. August 1983 - May 1989. Research conducted at the University of Texas Marine Science Institute, Port Aransas, Texas. Dissertation: Variable settlement of planktonic larvae in a marine benthic population: Spirorbis spirillum (Polychaeta: Spirorbidae) settling on blades of the seagrass Thalassia testudinum.
M.S. in Zoology. The University of Oklahoma, Norman, Oklahoma. August 1980 - July 1983. Fields of study: Ecology and aquatic invertebrate zoology. Thesis: The influence of wind mixing and river inflow on zooplankton vertical distribution and abundance in a large impoundment.
B.S. in Biology with co-major in Environmental Studies. Cum Laude. Baylor University, Waco, Texas. August 1976 - May 1980.