Ph.D. in Biological Sciences.  The University of Texas, Austin, Texas.  August 1983 - May 1989.  Research conducted at the University of Texas Marine Science Institute, Port Aransas, Texas.  Dissertation: Variable settlement of planktonic larvae in a marine benthic population:  Spirorbis spirillum (Polychaeta: Spirorbidae) settling on blades of the seagrass Thalassia testudinum.

 M.S. in Zoology.  The University of Oklahoma, Norman, Oklahoma.  August 1980 - July 1983.   Fields of study:  Ecology and aquatic invertebrate zoology.  Thesis:  The influence of wind mixing and river inflow on zooplankton vertical distribution and abundance in a large impoundment.

 B.S. in Biology with co-major in Environmental Studies.  Cum Laude.  Baylor University, Waco, Texas.  August 1976 - May 1980.